| # | Tower | No | Year | Materials | Maker | Trusses | Layout |
|---|
| 401 | Axbridge, Somerset, S John Bapt | 2 | | | | | |
| 402 | Axbridge, Somerset, S John Bapt | 1 | 1925 | Cast iron | John Taylor & Co | 8.3.C.e | 6.1 |
| 403 | Axminster, Devon, S Mary | 8 | | | | | |
| 404 | Axminster, Devon, S Mary | 2 | 1986 | Cast iron | Whitechapel Bell Foundry | 8.3.A.h | 3.1 |
| 405 | Axminster, Devon, S Mary | 1 | 1898† | Oak | Harry Stokes | 6.B | 8.3 |
| 406 | Axmouth, Devon, S Michael | 1 | c1699† | Oak | (unidentified) | 6.A | 4.1 |
| 407 | Aycliffe, Durham, S Andrew | 1 | 1931 | Metal | Gillett & Johnston | | |
| 408 | Aylburton, Gloucestershire, S Mary | 1 | | Oak | | 5.L | 1 |
| 409 | Aylesbeare, Devon, Blessed Virgin Mary | 2 | 1919 | Cast iron | John Taylor & Co | 8.3.A.h | |
| 410 | Aylesbeare, Devon, Blessed Virgin Mary | 1 | 1884 | Oak | Harry Stokes | 6.B | 5.1 |
| 411 | Aylesbury, Buckinghamshire, S Mary | 1 | 1983 | Cast iron | Eayre & Smith | 8.3.A.m | 8.3 |
| 412 | Aylesford, Kent, S Peter & S Paul | 1 | 1885 | Timber | Gillett & Co | 6.A | 8.3 |
| 413 | Aylestone, Leicestershire, S Andrew | 1 | 1948 | Cast iron | Gillett & Johnston | 8.3.A.m | 8.3 |
| 414 | Aylmerton, Norfolk, S John Bapt | 1 | | Timber | | 6.D | 1 |
| 415 | Aylsham, Norfolk, S Michael | 1 | 1889 | Oak | George Day & Sons | 6.D | 10.X |
| 416 | Aylsham, Norfolk, S Michael | 1.1 | | Oak | | | |
| 417 | Aymestrey, Herefordshire, S John Bapt & S Alkmund | 1 | c1699 | Oak | | 6.A | 6.16 |
| 418 | Aynho, Northamptonshire, S Michael | 2 | 1911 | Metal | John Taylor & Co | | |
| 419 | Aynho, Northamptonshire, S Michael | 1 | 1911 | Metal | John Taylor & Co | | |
| 420 | Aysgarth, North Yorkshire, S Andrew | 1 | 1913 | Cast iron | Gillett & Johnston | 8.3.C | 6 |
| 421 | Ayston, Rutland, S Mary V | 1 | 1877 | Oak | John Taylor & Co | | |
| 422 | Aythorpe Roding, Essex, S Mary | 1 | c1600† | Oak | | ~ 6.H | |
| 423 | Babbacombe, Devon, All Saints | 2 | 1963 | Cast iron | John Taylor & Co | 8.3.D.a | |
| 424 | Babbacombe, Devon, All Saints | 1 | 1963 | Cast iron | John Taylor & Co | 8.3.D.a | |
| 425 | Babcary, Somerset, Holy Cross | 1 | 1911 | Cast iron | John Warner & Sons | | 6 |
| 426 | Babraham, Cambridgeshire, S Peter | 1 | c1500 | Timber | | 5.H, 5.R & 6.D | 4.2 |
| 427 | Babworth, Nottinghamshire, All Saints | 1 | 1959 | Cast iron | John Taylor & Co | 8.3.A.h | 6.14 R |
| 428 | Backwell, Somerset, S Andrew | 9 | | Stone | | 9.A | 1 |
| 429 | Backwell, Somerset, S Andrew | 2 | 2018 | Cast iron | John Taylor & Co | 8.3.D.a | 3.X |
| 430 | Backwell, Somerset, S Andrew | 1 | 2018 | Cast iron | John Taylor & Co | 8.3.D.a | 7.X |
| 431 | Baconsthorpe, Norfolk, S Mary | 1 | | Oak | | 5.A | 1 |
| 432 | Bacton, Herefordshire, S Faith | 1 | 1907 | Oak | George Day & Sons | | 6 |
| 433 | Bacton, Suffolk, S Mary V | 1 | 1972 | Cast iron | Whitechapel Bell Foundry | | 6.4 |
| 434 | Bacup, Lancashire, Christ Church | 1 | c1854 | Timber | ? C & G Mears | 6.A | 6.1 ➔ 8.3 |
| 435 | Bacup, Lancashire, Christ Church | 1.1 | 1923 | Cast iron | ? Mears & Stainbank | 8.3.A.h | 2.X2 |
| 436 | Badby, Northamptonshire, S Mary | 1 | 1931 | Cast iron | John Taylor & Co | 8.3.A.h | 6.1 |
| 437 | Baddesley Clinton, Warwickshire, S Michael | 1 | | | | | |
| 438 | Baddesley Ensor, Warwickshire, S Nicholas | 1 | | | | | |
| 439 | Baddesley Ensor, Warwickshire, S Nicholas | 1.1 | | | | | |
| 440 | Badgeworth, Gloucestershire, Holy Trinity | 1 | 2011 | Steel | Nicholson Engineering Ltd | | 6.1 |
| 441 | Badgworth, Somerset, S Congar | 1 | c1748 | Oak | ? George Nott | 6.A | 5.1 |
| 442 | Badingham, Suffolk, S John Bapt | 1 | 1938 | Oak | Alfred Bowell | 6.A | 6.1 |
| 443 | Badley, Suffolk, S Mary | 1 | c1530 | Oak | | 6.H | 3.1 |
| 444 | Badminton, Gloucestershire, S Michael & All Angels | 2 | 1914 | Cast iron | Gillett & Johnston | 8.3.D.a | 1 |
| 445 | Badminton, Gloucestershire, S Michael & All Angels | 1 | 1914 | Cast iron | Gillett & Johnston | 8.3.D.a | 5.1 |
| 446 | Badsey, Worcestershire, S James | 1 | 1897 | Cast iron | John Taylor & Co | 8.3.A.h | 8.3 |
| 447 | Badshot Lea, Surrey, S George | 1 | | Stone | | 9.A | 1 |
| 448 | Badsworth, West Yorkshire, S Mary V | 1 | 1916 | Cast iron | John Taylor & Co | 8.3.A.h | 6.1 |
| 449 | Badwell Ash, Suffolk, S Mary | 1 | c1499 | Oak | | 5.G & ~ 5.K | 5.12 R |
| 450 | Bagendon, Gloucestershire, S Margaret | 1 | 1897 | Timber | Gillett & Johnston | | |